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The Carboniferous Portal

Introduction

The Carboniferous ( /ˌkɑːrbəˈnɪfərəs/ KAR-bə-NIF-ər-əs) is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period 358.9 Ma (million years ago) to the beginning of the Permian Period, 298.9 Ma. In North America, the Carboniferous is often treated as two separate geological periods, the earlier Mississippian and the later Pennsylvanian.

The name Carboniferous means " coal-bearing", from the Latin carbō (" coal") and ferō ("bear, carry"), and refers to the many coal beds formed globally during that time. The first of the modern "system" names, it was coined by geologists William Conybeare and William Phillips in 1822, based on a study of the British rock succession.

Carboniferous is the period during which both terrestrial animal and land plant life was well established. Stegocephalia (four-limbed vertebrates including true tetrapods), whose forerunners ( tetrapodomorphs) had evolved from lobe-finned fish during the preceding Devonian period, became pentadactylous during the Carboniferous. The period is sometimes called the Age of Amphibians because of the diversification of early amphibians such as the temnospondyls, which became dominant land vertebrates, as well as the first appearance of amniotes including synapsids (the clade to which modern mammals belong) and sauropsids (which include modern reptiles and birds) during the late Carboniferous. Due to the raised atmospheric oxygen level, land arthropods such as arachnids (e.g. trigonotarbids and Pulmonoscorpius), myriapods (e.g. Arthropleura) and insects (e.g. Meganeura) also underwent a major evolutionary radiation during the late Carboniferous. Vast swaths of forests and swamps covered the land, which eventually became the coal beds characteristic of the Carboniferous stratigraphy evident today.

The later half of the period experienced glaciations, low sea level, and mountain building as the continents collided to form Pangaea. A minor marine and terrestrial extinction event, the Carboniferous rainforest collapse, occurred at the end of the period, caused by climate change. ( Full article...)

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Arthropods are members of the phylum Arthropoda, and include the insects, arachnids, and crustaceans. They are characterized by their jointed limbs and cuticles. The rigid cuticle inhibits growth, so arthropods replace it periodically by moulting. The arthropod body plan consists of repeated segments, each with a pair of appendages. Their versatility has enabled them to become the most species-rich members of all ecological guilds in most environments. They have over a million described species, making up more than 80% of all described living animal species. They range in size from microscopic plankton up to forms a few meters long.

Like their exteriors, the internal organs of arthropods are generally built of repeated segments. Their vision relies on various combinations of compound eyes and pigment-pit ocelli. Arthropods also have a wide range of chemical and mechanical sensors, mostly based on modifications of the many setae (bristles) that project through their cuticles. Nearly all arthropods lay eggs. Arthropod hatchlings vary from miniature adults to grubs and caterpillars that lack jointed limbs and eventually undergo a total metamorphosis to produce the adult form.

The evolutionary ancestry of arthropods dates back to the Cambrian period. The group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians (or their constituent clades) in a superphylum Ecdysozoa. Overall however, the basal relationships of Metazoa are not yet well resolved. Likewise, the relationships between various arthropod groups are still actively debated. ( see more...)

Did you know...

... that siltstone deposits have allowed three-dimensional fossils of the extinct mushroom crinoid to be excavated from Indiana?
...that eighty years on, scientists are still debating whether the Palæozoic fossils known as Chitinozoans ( SEM image pictured) represent plants, animals or eggs?
... that Arthropleuridea is an extinct class of myriapods which includes, at over 2 meters long, the largest terrestrial arthropods that ever lived?

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Eli Heimans' reconstruction (1911) of the Carboniferous tree Lepidodendron, known from fossil " Stigmaria" (fossil roots) in Limburg. Drawing also shows a piece of fossil bast.
Photo credit: Eli Heimans.

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The history of paleontology traces the history of the effort to study the fossil record left behind by ancient life forms. Although fossils had been studied by scholars since ancient times, the nature of fossils and their relationship to life in the past became better understood during the 17th and 18th centuries. At the end of the 18th century the work of Georges Cuvier ended a long running debate about the reality of extinction and led to the emergence of paleontology as a scientific discipline.

The first half of the 19th century saw paleontological activity become increasingly well organized. This contributed to a rapid increase in knowledge about the history of life on Earth, and progress towards definition of the geologic time scale. As knowledge of life's history continued to improve, it became increasingly obvious that there had been some kind of successive order to the development of life. After Charles Darwin published Origin of Species in 1859, much of the focus of paleontology shifted to understanding evolutionary paths.

The last half of the 19th century saw a tremendous expansion in paleontological activity, especially in North America. The trend continued in the 20th century with additional regions of the Earth being opened to systematic fossil collection, as demonstrated by a series of important discoveries in China near the end of the 20th century. There was also a renewed interest in the Cambrian explosion that saw the development of the body plans of most animal phyla. ( see more...)