From Wikipedia, the free encyclopedia
Genus of beetles
Harpalus is a
genus of
ground beetle first described by
Pierre André Latreille in 1802.
[1]
Harpalus contains the following 430 species:
[2]
[3]
- Subgenus Cephalophonus Ganglbauer, 1891
-
Harpalus cephalotes Fairmaire & Laboulbène, 1854
- Subgenus Cryptophonus Brandmayr & Zetto Brandmayr, 1982
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Harpalus agnatus Reiche, 1850
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Harpalus cyrenaicus Koch, 1939
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Harpalus fulvus Dejean, 1829
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Harpalus grilli Kataev, 2002
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Harpalus idiotus Bates, 1889
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Harpalus janinae Jeanne, 1984
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Harpalus litigiosus Dejean, 1829
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Harpalus melancholicus Dejean, 1829
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Harpalus schaumii Wollaston, 1864
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Harpalus tenebrosus Dejean, 1829
- Subgenus Glanodes Casey, 1914
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Harpalus cohni Ball, 1972
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Harpalus corpulentus (Casey, 1914)
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Harpalus huachuca Ball, 1972
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Harpalus obliquus G.Horn, 1880
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Harpalus puncticeps (Casey, 1914)
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Harpalus stephani Ball, 1972
- Subgenus Harpalus Latreille, 1802
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Harpalus acupalpoides Reitter, 1900
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Harpalus aeneipennis (Faldermann, 1836)
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Harpalus aequicollis Motschulsky, 1844
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Harpalus affinis (Schrank, 1781)
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Harpalus agakhaniantzi (Mikhailov, 1972)
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Harpalus akbensis Jedlicka, 1958
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Harpalus akinini Tschitscherine, 1895
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Harpalus alajensis Tschitscherine, 1898
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Harpalus albanicus Reitter, 1900
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Harpalus alexandrae Kataev, 1990
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Harpalus alexeevi Kataev, 1990
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Harpalus alpivagus Tschitscherine, 1899
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Harpalus amarellus Bates, 1891
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Harpalus amariformis Motschulsky, 1844
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Harpalus amplicollis Ménétriés, 1848
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Harpalus amputatus Say, 1830
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Harpalus anatolicus Tschitscherine, 1898
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Harpalus angulatus Putzeys, 1878
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Harpalus angustipennis Boheman, 1848
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Harpalus angustitarsis Reitter, 1887
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Harpalus animosus Casey, 1924
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Harpalus anisodactyliformis Solsky, 1874
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Harpalus antonowi (Tschitscherine, 1898)
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Harpalus anxioides Kataev, 1991
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Harpalus anxius (Duftschmid, 1812)
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Harpalus araraticus Mlynar, 1979
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Harpalus arcuatus Tschitscherine, 1898
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Harpalus arnoldii Kataev, 1988
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Harpalus ascetes Kataev & Wrase, 1997
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Harpalus asemus Basilewsky, 1946
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Harpalus aterrimus Casey, 1914
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Harpalus atratus Latreille, 1804
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Harpalus atrichatus Hatch, 1949
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Harpalus atripes Casey, 1914
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Harpalus attenuatus Stephens, 1828
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Harpalus australasiae Dejean, 1829
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Harpalus autumnalis (Duftschmid, 1812)
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Harpalus badakschanus Jedlicka, 1956
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Harpalus baleensis Clarke, 1973
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Harpalus balli Noonan, 1991
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Harpalus basanicus J.Sahlberg, 1913
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Harpalus basilewskyi Facchini, 2003
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Harpalus basuto Basilewsky, 1958
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Harpalus bellieri Reiche, 1861
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Harpalus beneshi Kataev & Wrase, 1997
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Harpalus brachypterus Tschitscherine, 1898
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Harpalus brevicornis Germar, 1823
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Harpalus brevis Motschulsky, 1844
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Harpalus breviusculus Chaudoir, 1846
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Harpalus bucharicus Tschitscherine, 1898
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Harpalus bulirschi (Facchini, 2012)
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Harpalus bungii Chaudoir, 1844
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Harpalus caeruleatus Bates, 1878
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Harpalus caiphus Reiche & Saulcy, 1855
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Harpalus calathoides Motschulsky, 1844
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Harpalus capicola Dejean, 1829
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Harpalus cardoni Antoine, 1922
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Harpalus caspius (Steven, 1806)
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Harpalus cautus Dejean, 1829
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Harpalus chalcentus Bates, 1873
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Harpalus chasanensis Lafer, 1989
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Harpalus chobautianus Lutshnik, 1922
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Harpalus chrysopus Reitter, 1887
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Harpalus circumpunctatus Chaudoir, 1846
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Harpalus cisteloides Motschulsky, 1844
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Harpalus clarkei Kataev & J.Schmidt, 2020
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Harpalus compressus Motschulsky, 1844
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Harpalus contemptus Dejean, 1829
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Harpalus convexus Faldermann, 1836
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Harpalus corporosus (Motschulsky, 1862)
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Harpalus corrugatus Basilewsky, 1958
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Harpalus crates Bates, 1883
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Harpalus cupreus Dejean, 1829
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Harpalus cursorius Péringuey, 1896
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Harpalus cyanopterus (Tschitscherine, 1897)
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Harpalus cyclogonus Chaudoir, 1844
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Harpalus danieli Reitter, 1900
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Harpalus davidianus Tschitscherine in Jakobson, 1903
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Harpalus decipiens Dejean, 1829
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Harpalus defector Péringuey, 1896
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Harpalus desertus LeConte, 1859
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Harpalus diligens Tschitscherine, 1898
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Harpalus dimidiatus (P.Rossi, 1790)
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Harpalus dispar Dejean, 1829
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Harpalus dissitus Antoine, 1931
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Harpalus distinguendus (Duftschmid, 1812)
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Harpalus diversicollis Basilewsky, 1958
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Harpalus dubius Boheman, 1848
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Harpalus dudkoi Kataev, 2011
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Harpalus durangoensis Bates, 1891
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Harpalus ebeninus Heyden, 1870
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Harpalus egenus Dejean, 1829
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Harpalus egorovi Lafer, 1989
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Harpalus ellipsis LeConte, 1847
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Harpalus elliptipennis Facchini, 2003
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Harpalus erosus Mannerheim, 1825
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Harpalus exiguus Boheman, 1848
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Harpalus famelicus Tschitscherine, 1898
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Harpalus farkaci Kataev & Wrase, 1995
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Harpalus fimetarius Dejean, 1829
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Harpalus flavescens (Piller & Mitterpacher, 1783)
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Harpalus flavicornis Dejean, 1829
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Harpalus foveiger Tschitscherine, 1895
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Harpalus franzi Mateu, 1954
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Harpalus frater Chaudoir, 1876
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Harpalus fraternus LeConte, 1852
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Harpalus froelichii Sturm, 1818
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Harpalus fulvicornis Thunberg, 1806
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Harpalus fulvilabris Mannerheim, 1853
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Harpalus fulvipennis Chaudoir, 1843
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Harpalus fuscicornis Ménétriés, 1832
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Harpalus fuscipalpis Sturm, 1818
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Harpalus fuscipennis Wiedemann, 1825
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Harpalus fuscoaeneus Dejean, 1829
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Harpalus ganssuensis Semenov, 1889
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Harpalus giacomazzoi Kataev & Wrase, 1996
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Harpalus gilgil Basilewsky, 1946
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Harpalus gisellae Csiki, 1932
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Harpalus glasunovi Kataev, 1987
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Harpalus gravis LeConte, 1858
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Harpalus gregoryi Alluaud, 1917
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Harpalus hartmanni Kataev, 2002
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Harpalus herbivagus Say, 1823
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Harpalus heyrovskyi Jedlicka, 1928
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Harpalus hiekei Kataev & Wrase, 2010
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Harpalus hirtipes (Panzer, 1796)
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Harpalus honestus (Duftschmid, 1812)
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Harpalus hospes Sturm, 1818
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Harpalus hybridus Boheman, 1848
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Harpalus impressus Roth, 1851
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Harpalus inconcinnus Chaudoir, 1876
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Harpalus indianus Csiki, 1932
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Harpalus indicola Bates, 1878
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Harpalus indigens Casey, 1924
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Harpalus inexspectatus Kataev, 1989
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Harpalus ingenuus Tschitscherine, 1898
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Harpalus innocuus LeConte, 1863
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Harpalus italus Schaum, 1860
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Harpalus jeanneli Basilewsky, 1946
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Harpalus jordanus Jedlicka, 1964
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Harpalus kabakianus Kataev, 1988
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Harpalus kabakovi Kataev, 1987
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Harpalus kadleci Kataev & Wrase, 1995
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Harpalus kadyrbekovi Kataev, 1988
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Harpalus kagyzmanicus Kataev, 1984
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Harpalus kandaharensis Jedlicka, 1956
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Harpalus karakorum Jedlicka, 1958
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Harpalus karamani Apfelbeck, 1902
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Harpalus kazanensis Jedlicka, 1958
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Harpalus kaznakovi Kataev & Wrase, 1997
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Harpalus kibonoti Alluaud, 1927
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Harpalus kirgisicus (Motschulsky, 1844)
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Harpalus kiritshenkoi Kataev, 1990
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Harpalus klapperichi Jedlicka, 1956
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Harpalus kmecoi Facchini, 2003
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Harpalus kozlovi Kataev, 1993
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Harpalus kryzhanovskii Kataev, 1988
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Harpalus kunarensis Kataev, 1993
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Harpalus laetus Reiche, 1843
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Harpalus laevipes Zetterstedt, 1828
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Harpalus lama Kataev & Wrase, 1997
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Harpalus lateralis Dejean, 1829
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Harpalus laticeps LeConte, 1850
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Harpalus latus (Linnaeus, 1758)
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Harpalus lederi Tschitscherine, 1899
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Harpalus lethierryi Reiche, 1860
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Harpalus lewisii LeConte, 1865
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Harpalus longipalmatus Mordkovitsh, 1969
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Harpalus lopezi J.Serrano & Lencina, 2009
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Harpalus lugubris Boheman, 1848
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Harpalus lumbaris Mannerheim, 1825
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Harpalus luteicornis (Duftschmid, 1812)
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Harpalus lutshniki Schauberger, 1932
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Harpalus macronotus Tschitscherine, 1893
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Harpalus mairei Peyerimhoff, 1928
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Harpalus major (Motschulsky, 1850)
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Harpalus makhekensis Basilewsky, 1958
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Harpalus manas Kataev, 1990
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Harpalus marginellus Gyllenhal, 1827
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Harpalus martini Van Dyke, 1926
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Harpalus masoreoides Bates, 1878
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Harpalus massarti Burgeon, 1936
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Harpalus mauritanicus Gaubil, 1844
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Harpalus medvedevi Kataev, 2006
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Harpalus megacephalus LeConte, 1847
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Harpalus melaneus Bates, 1878
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Harpalus metallinus Ménétriés, 1836
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Harpalus metarsius Andrewes, 1930
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Harpalus meteorus Basilewsky, 1946
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Harpalus michaili Kataev, 1990
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Harpalus michailovi Kataev, 1987
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Harpalus microthorax (Motschulsky, 1849)
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Harpalus miles Péringuey, 1896
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Harpalus minutissimus Facchini, 2015
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Harpalus minutulus Kataev & Liang, 2004
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Harpalus mitridati Pliginskiy, 1915
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Harpalus mlynari Kataev, 1990
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Harpalus modestus Dejean, 1829
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Harpalus morvani Kataev, 2002
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Harpalus murzini Kataev, 2008
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Harpalus nanniscus Péringuey, 1896
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Harpalus natalensis Boheman, 1848
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Harpalus natalicus Péringuey, 1896
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Harpalus neglectus Audinet-Serville, 1821
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Harpalus nevadensis K. & J.Daniel, 1898
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Harpalus nigrans A.Morawitz, 1862
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Harpalus nigritarsis C.R.Sahlberg, 1827
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Harpalus numidicus Bedel, 1893
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Harpalus nyassicus Basilewsky, 1946
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Harpalus oblitus Dejean, 1829
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Harpalus obnixus Casey, 1924
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Harpalus ochropus Kirby, 1837
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Harpalus oodioides Dejean, 1829
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Harpalus opacipennis (Haldeman, 1843)
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Harpalus optabilis Dejean, 1829
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Harpalus ovtshinnikovi Kataev, 1990
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Harpalus pallidipennis A.Morawitz, 1862
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Harpalus pallipes Chaudoir, 1837
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Harpalus parallelocollis Facchini, 2003
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Harpalus parasinuatus Kataev & Liang, 2007
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Harpalus parvulus Dejean, 1829
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Harpalus petri Tschitscherine, 1902
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Harpalus picipennis (Duftschmid, 1812)
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Harpalus plancyi Tschitscherine, 1897
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Harpalus plenalis Casey, 1914
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Harpalus politus Dejean, 1829
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Harpalus polyglyptus Schaum, 1862
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Harpalus potanini Tschitscherine, 1906
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Harpalus praecurrens Schauberger, 1934
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Harpalus procognatus Lorenz, 1998
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Harpalus progrediens Schauberger, 1922
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Harpalus prosperus Basilewsky, 1972
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Harpalus providens Casey, 1914
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Harpalus pseudoserripes Reitter, 1900
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Harpalus pterostichus (Reitter, 1900)
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Harpalus puetzi Kataev & Wrase, 1997
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Harpalus pulchrinulus Reitter, 1900
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Harpalus pulvinatus Ménétriés, 1848
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Harpalus pumilus Sturm, 1818
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Harpalus punctatostriatus Dejean, 1829
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Harpalus punctipennis Mulsant, 1852
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Harpalus pusillus (Motschulsky, 1850)
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Harpalus pygmaeus Dejean, 1829
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Harpalus quadratus Chaudoir, 1846
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Harpalus raphaili Kataev, 1997
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Harpalus reflexus Putzeys in Schneider & Leder, 1878
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Harpalus reitteri Wrase & Kataev, 2011
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Harpalus remboides Solsky, 1874
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Harpalus reversus Casey, 1924
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Harpalus rhemboides Solsky, 1874
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Harpalus rivalsi Jeannel, 1948
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Harpalus rotundus Facchini, 2003
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Harpalus rougemonti Clarke, 1973
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Harpalus rubripes (Duftschmid, 1812)
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Harpalus rufipalpis Sturm, 1818
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Harpalus rufiscapus Gebler, 1833
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Harpalus rufitarsoides Schauberger, 1934
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Harpalus rufomarginatus (Boheman, 1848)
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Harpalus rumelicus Apfelbeck, 1904
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Harpalus salinulus Reitter, 1900
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Harpalus salinus Dejean, 1829
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Harpalus sanctaehelenae Basilewsky, 1972
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Harpalus saxicola Dejean, 1829
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Harpalus semenowi Tschitscherine, 1901
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Harpalus semipunctatus Dejean, 1829
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Harpalus serripes (Quensel, 1806)
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Harpalus servus (Duftschmid, 1812)
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Harpalus setiporus (Reitter, 1894)
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Harpalus siculus Dejean, 1829
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Harpalus silipes Dejean, 1831
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Harpalus sinuaticollis Facchini, 2015
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Harpalus sinuatus Tschitscherine, 1893
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Harpalus smaragdinus (Duftschmid, 1812)
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Harpalus smyrnensis Heyden, 1888
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Harpalus solitaris Dejean, 1829
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Harpalus somereni Basilewsky, 1946
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Harpalus somnulentus Dejean, 1829
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Harpalus spadiceus Dejean, 1829
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Harpalus spretus Péringuey, 1896
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Harpalus spurius Péringuey, 1896
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Harpalus stevenii Dejean, 1829
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Harpalus stevensi Kataev, 2011
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Harpalus stoetznerianus Schauberger, 1932
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Harpalus strenuus Tschitscherine, 1898
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Harpalus subaeneus Boheman, 1848
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Harpalus subcylindricus Dejean, 1829
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Harpalus subtruncatus Chaudoir, 1846
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Harpalus sulphuripes Germar, 1823
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Harpalus sundaicus Schauberger, 1933
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Harpalus sushenicus Kataev, 1990
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Harpalus suturangulus Reitter, 1887
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Harpalus szalliesi Kataev & Wrase, 1995
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Harpalus taciturnus Dejean, 1829
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Harpalus tangutorum Kataev, 1993
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Harpalus tardus (Panzer, 1796)
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Harpalus tarsalis Mannerheim, 1825
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Harpalus terrestris (Motschulsky, 1844)
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Harpalus tibeticus Andrewes, 1930
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Harpalus tichonis Jakobson, 1907
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Harpalus tiridates Reitter, 1900
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Harpalus tithonus Reitter, 1900
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Harpalus tjanschanicus Semenov, 1889
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Harpalus torosensis Jedlicka, 1961
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Harpalus torridoides Reitter, 1900
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Harpalus trichophorus Tschitscherine, 1897
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Harpalus triseriatus A.Fleischer, 1897
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Harpalus turcicus Jedlicka, 1958
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Harpalus turmalinus Erichson, 1847
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Harpalus udege Lafer, 1989
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Harpalus uniformis (Motschulsky, 1844)
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Harpalus ussuricus Mlynar, 1979
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Harpalus vanemdeni Schauberger, 1932
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Harpalus venator Boheman, 1848
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Harpalus ventralis LeConte, 1847
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Harpalus vereschaginae Kataev, 1988
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Harpalus vernicosus Kataev & Liang, 2007
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Harpalus viridanus (Motschulsky, 1844)
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Harpalus viridicupreus Reiche, 1843
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Harpalus vittatus Gebler, 1833
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Harpalus wadiensis Jedlicka, 1964
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Harpalus wohlberedti Emden, 1932
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Harpalus xanthopus Gemminger & Harold, 1868
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Harpalus zabroides Dejean, 1829
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Harpalus zhdankoi Kataev, 1990
- Subgenus Loboharpalus Schauberger, 1932
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Harpalus platynotus Bates, 1873
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Harpalus rubefactus Bates, 1873
- Subgenus Opadius Casey, 1914
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Harpalus apache Kataev, 2010
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Harpalus cordatus (LeConte, 1853)
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Harpalus cordifer Notman, 1919
- Subgenus Plectralidus Casey, 1914
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Harpalus erraticus Say, 1823
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Harpalus retractus LeConte, 1863
- Subgenus Pseudoophonus Motschulsky, 1844
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Harpalus actiosus Casey, 1914
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Harpalus aenigma (Tschitscherine, 1897)
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Harpalus alienus Bates, 1878
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Harpalus aogashimensis (Habu, 1957)
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Harpalus azumai Habu, 1968
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Harpalus babai Habu, 1973
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Harpalus calceatus (Duftschmid, 1812)
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Harpalus caliginosus (Fabricius, 1775)
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Harpalus capito A.Morawitz, 1862
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Harpalus compar LeConte, 1847
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Harpalus coreanus (Tschitscherine, 1895)
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Harpalus davidi (Tschitscherine, 1897)
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Harpalus disimuciulus Huang; Lei; Yan & Hu, 1996
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Harpalus eous Tschitscherine, 1901
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Harpalus erythropus Dejean, 1829
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Harpalus faunus Say, 1823
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Harpalus fokienensis Schauberger, 1930
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Harpalus griseus (Panzer, 1796)
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Harpalus hatchi Ball & Anderson, 1962
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Harpalus hauserianus Schauberger, 1929
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Harpalus indicus Bates, 1891
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Harpalus jureceki (Jedlicka, 1928)
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Harpalus katiae F.Battoni, 1985
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Harpalus liobasis Chaudoir, 1868
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Harpalus meghalayensis Kataev, 2001
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Harpalus meridianus Andrewes, 1923
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Harpalus paratus Casey, 1924
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Harpalus pastor Motschulsky, 1844
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Harpalus pensylvanicus (DeGeer, 1774)
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Harpalus poncei Will, 2002
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Harpalus protractus Casey, 1914
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Harpalus pseudohauserianus Kataev, 2001
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Harpalus pseudophonoides Schauberger, 1930
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Harpalus roninus Bates, 1873
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Harpalus rufipes (DeGeer, 1774)
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Harpalus sericatus Tschitscherine, 1906
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Harpalus simplicidens Schauberger, 1929
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Harpalus singularis Tschitscherine, 1906
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Harpalus sinicus Hope, 1845
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Harpalus suensoni Kataev, 1997
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Harpalus texanus Casey, 1914
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Harpalus tridens A.Morawitz, 1862
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Harpalus ussuriensis Chaudoir, 1863
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Harpalus vagans LeConte, 1865
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Harpalus yinchuanensis Huang, 1993
- Subgenus Semiophonus Schauberger, 1933
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Harpalus signaticornis (Duftschmid, 1812)
- Subgenus Zangoharpalus Huang, 1998
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Harpalus mariae Kataev, 1997
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Harpalus microdemas Schauberger, 1932
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Harpalus praticola Bates, 1891
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Harpalus pseudotinctulus Schauberger, 1932
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Harpalus tinctulus Bates, 1873
- Extinct, not assigned to subgenus
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Harpalus abolitus C. & L.Heyden, 1866
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Harpalus anactus Giebel, 1856
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Harpalus bruckmanni Heer, 1862
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Harpalus burmeisteri Giebel, 1856
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Harpalus conditus Scudder, 1900
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Harpalus constrictus Heer, 1862
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Harpalus deletus Oustalet, 1864
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Harpalus dilatatus Theobald, 1937
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Harpalus ewaldi Giebel, 1856
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Harpalus excavatus Forster, 1891
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Harpalus heeri Giebel, 1856
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Harpalus knorri Giebel, 1856
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Harpalus lheritieri Piton, 1940
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Harpalus liasinus Giebel, 1856
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Harpalus maceratus Wickham, 1911
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Harpalus nero Oustalet, 1864
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Harpalus nitens Birket-Smith, 1977
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Harpalus nuperus Scudder, 1900
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Harpalus obtusus Forster, 1891
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Harpalus oustaleti Meunier, 1915
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Harpalus pleistocenicus Lomnicki, 1894
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Harpalus redivivus Wickham, 1917
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Harpalus schlotheimi Giebel, 1856
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Harpalus sinis Heer, 1847
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Harpalus stierlini Heer, 1862
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Harpalus stygius Heer, 1862
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Harpalus tabidus Heer, 1847
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Harpalus tardigradus Heer, 1862
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Harpalus ulomaeformis Wickham, 1917
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Harpalus veterum Cockerell, 1920
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Harpalus whitfieldi Scudder, 1900