Dihoplus Temporal range:
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Dihoplus schleiermacheri holotype skull | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Family: | Rhinocerotidae |
Tribe: | Dicerorhinini |
Genus: | †
Dihoplus Brandt, 1878 |
Type species | |
†Rhinoceros schleiermacheri Kaup, 1832
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Species | |
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Dihoplus is an extinct genus of rhinoceros that lived in Eurasia from the Late Miocene to Pliocene. [1]
Species of Dihoplus were large rhinoceroses, with the body masses of Dihoplus schleiermacheri and Dihoplus pikermiensis estimated at 2,122 kilograms (4,678 lb) and 1,100 kilograms (2,400 lb) respectively. [2] The head bore two horns. The nasal septum was not ossified, with a nasal notch above the frontmost premolars. The toothrow is placed posteriorly within the skull. The first upper premolar is absent, though the lower second incisor is present. [1]
Members of Dihoplus were long placed in Dicerorhinus (which contains the living Sumatran rhinoceros). Sometimes these species are placed in the related Stephanorhinus. The genus is now generally considered distinct, [3] though there is still debate as to which species should be included; for example, Deng (2011) listed Merck's rhinoceros ( Stephanorhinus kirchbergensis) under Dihoplus. [4] Species recently placed in the genus include: [5]
"D." megarhinus (de Christol, 1834) from the Late Miocene-Pliocene of Europe, Anatolia, China and Transbaikalia, [1] [5] since 2021 has been placed instead in the genus Pliorhinus along with "Dicerorhinus" miguelcrusafonti from the Pliocene of Europe. [6] [7] "D." ringstoemi is either considered a synonym of "D." megarhinus [1] or placed as a separate species within Pliorhinus. [8]
The monophyly of the genus has been questioned, with some studies suggesting that D. pikermiensis is more closely related to Stephanorhinus and Coelodonta (which contains the woolly rhinoceros) than to the type species D. schleiermacheri. [7]
Morphological phylogeny after Pandolfi (2023), excluding living African rhinoceros species. [7]
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D. pikermiensis and D. bethlehemsis are suggested to have been browsers. [9] [5]
Dihoplus Temporal range:
| |
---|---|
Dihoplus schleiermacheri holotype skull | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Family: | Rhinocerotidae |
Tribe: | Dicerorhinini |
Genus: | †
Dihoplus Brandt, 1878 |
Type species | |
†Rhinoceros schleiermacheri Kaup, 1832
| |
Species | |
|
Dihoplus is an extinct genus of rhinoceros that lived in Eurasia from the Late Miocene to Pliocene. [1]
Species of Dihoplus were large rhinoceroses, with the body masses of Dihoplus schleiermacheri and Dihoplus pikermiensis estimated at 2,122 kilograms (4,678 lb) and 1,100 kilograms (2,400 lb) respectively. [2] The head bore two horns. The nasal septum was not ossified, with a nasal notch above the frontmost premolars. The toothrow is placed posteriorly within the skull. The first upper premolar is absent, though the lower second incisor is present. [1]
Members of Dihoplus were long placed in Dicerorhinus (which contains the living Sumatran rhinoceros). Sometimes these species are placed in the related Stephanorhinus. The genus is now generally considered distinct, [3] though there is still debate as to which species should be included; for example, Deng (2011) listed Merck's rhinoceros ( Stephanorhinus kirchbergensis) under Dihoplus. [4] Species recently placed in the genus include: [5]
"D." megarhinus (de Christol, 1834) from the Late Miocene-Pliocene of Europe, Anatolia, China and Transbaikalia, [1] [5] since 2021 has been placed instead in the genus Pliorhinus along with "Dicerorhinus" miguelcrusafonti from the Pliocene of Europe. [6] [7] "D." ringstoemi is either considered a synonym of "D." megarhinus [1] or placed as a separate species within Pliorhinus. [8]
The monophyly of the genus has been questioned, with some studies suggesting that D. pikermiensis is more closely related to Stephanorhinus and Coelodonta (which contains the woolly rhinoceros) than to the type species D. schleiermacheri. [7]
Morphological phylogeny after Pandolfi (2023), excluding living African rhinoceros species. [7]
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D. pikermiensis and D. bethlehemsis are suggested to have been browsers. [9] [5]